What happens to volatile keys when loading from RDB in Redis?

redis rdb ttl

foo@bar:~$ redis-cli 
127.0.0.1:6379> flushall
OK
127.0.0.1:6379> save
OK
127.0.0.1:6379> set p persistent
OK
127.0.0.1:6379> set v volatile
OK
127.0.0.1:6379> expire v 99999999
(integer) 1
127.0.0.1:6379> ttl v
(integer) 99999997
127.0.0.1:6379> save
OK
127.0.0.1:6379> shutdown
not connected> exit
foo@bar:~$ sudo service redis_6379 start
[sudo] password for foo: 
Starting Redis server...
foo@bar:~$ redis-cli 
127.0.0.1:6379> dbsize
(integer) 2
127.0.0.1:6379> ttl p
(integer) -1
127.0.0.1:6379> ttl v
(integer) 99999956