foo.

diff --git a/django/db/models/base.py b/django/db/models/base.py
index a0f43de..a6b5bb0 100644
--- a/django/db/models/base.py
+++ b/django/db/models/base.py
@@ -648,23 +648,25 @@ class Model(six.with_metaclass(ModelBase)):
         """
         meta = cls._meta
         for parent, field in meta.parents.items():
+            self._save_parents(cls=parent, using=using, update_fields=update_fields)
+
             # Make sure the link fields are synced between parent and self.
-            if (field and getattr(self, parent._meta.pk.attname) is None
-                    and getattr(self, field.attname) is not None):
+            if (getattr(self, field.attname) is not None):
                 setattr(self, parent._meta.pk.attname, getattr(self, field.attname))
-            self._save_parents(cls=parent, using=using, update_fields=update_fields)
+
             self._save_table(cls=parent, using=using, update_fields=update_fields)
+
             # Set the parent's PK value to self.
-            if field:
-                setattr(self, field.attname, self._get_pk_val(parent._meta))
-                # Since we didn't have an instance of the parent handy set
-                # attname directly, bypassing the descriptor. Invalidate
-                # the related object cache, in case it's been accidentally
-                # populated. A fresh instance will be re-built from the
-                # database if necessary.
-                cache_name = field.get_cache_name()
-                if hasattr(self, cache_name):
-                    delattr(self, cache_name)
+            setattr(self, field.attname, self._get_pk_val(parent._meta))
+
+            # Since we didn't have an instance of the parent handy set
+            # attname directly, bypassing the descriptor. Invalidate
+            # the related object cache, in case it's been accidentally
+            # populated. A fresh instance will be re-built from the
+            # database if necessary.
+            cache_name = field.get_cache_name()
+            if hasattr(self, cache_name):
+                delattr(self, cache_name)
 
     def _save_table(self, raw=False, cls=None, force_insert=False,
                     force_update=False, using=None, update_fields=None):