#14984 Link-to Model not Working

controllers.application.js

import Ember from 'ember';

export default Ember.Controller.extend({
  appName: 'Ember Twiddle'
});

// Broken on 2.10.2
// Works as expected on 2.9.1

models.client.js

import Model from "ember-data/model";
import attr from "ember-data/attr";
import { belongsTo, hasMany } from "ember-data/relationships";

export default Model.extend({
  name: attr('string'),
  invoices: hasMany('invoice')
});

models.invoice.js

import Model from "ember-data/model";
import attr from "ember-data/attr";
import { belongsTo, hasMany } from "ember-data/relationships";

export default Model.extend({
  number: attr('string'),
  client: belongsTo('client')
});

router.js

import Ember from 'ember';
import config from './config/environment';

const Router = Ember.Router.extend({
  location: 'none',
  rootURL: config.rootURL
});

Router.map(function() {
  this.route('single', {path: '/:id'});
});

export default Router;

routes.index.js

import Ember from 'ember';

export default Ember.Route.extend({
  model(){
      var client = this.store.createRecord('client', {name: 'Ross'});
      
      this.store.createRecord('invoice', {number: 'R0001', client: client});
			this.store.createRecord('invoice', {number: 'F0002', client: client});
      
      return this.store.peekAll('invoice');
  }
});

routes.single.js

import Ember from 'ember';

export default Ember.Route.extend({
});

templates.application.hbs

<h1>Welcome to {{appName}}</h1>
<br>
<br>
{{outlet}}
<br>
<br>

templates.index.hbs

<div class="row">
	{{#each model as |invoice|}}
		{{#link-to "single" invoice.client.id}}
    	{{invoice.number}}: {{invoice.client.name}}
    {{/link-to}}<br/>
	{{else}}
		No clients to show
	{{/each}}
</div>

twiddle.json

{
  "version": "0.15.1",
  "EmberENV": {
    "FEATURES": {}
  },
  "options": {
    "use_pods": false,
    "enable-testing": false
  },
  "dependencies": {
    "jquery": "https://cdnjs.cloudflare.com/ajax/libs/jquery/3.3.1/jquery.js",
    "ember": "3.4.3",
    "ember-template-compiler": "3.4.3",
    "ember-testing": "3.4.3"
  },
  "addons": {
    "ember-data": "3.4.2"
  }
}